john hawks weblog

paleoanthropology, genetics and evolution

distal

  • A quick start on anatomical directions

    Mon, 2013-01-21 23:57 -- John Hawks
    Synopsis: 
    A laboratory station helping to orient on directions in anatomy

    When talking about bones and teeth, we will need to use several terms to orient ourselves. Some of the terms are obvious, like right and left. Other intuitive terms can fail us, however. For example, we could use higher and lower to refer to parts of our arms, but these terms will be confusing if we lift our arms over our heads. Even left and right can cause confusion: sometimes we need to talk about the left surface of our right arm, for instance. For reasons like these, anthropologists use terms with specific anatomical meanings to talk about the
    positions of bones and features on them.

    Humans are special compared to many vertebrates in having a vertebral axis that runs roughly up and down, at least while we are standing up. For this reason, a long tradition in human anatomy uses these terms:

    Superior: Higher. The nose is superior to the mouth.

    Inferior: Lower. The nostrils are most visible on the inferior aspect of the nose.

    These terms are always used when referring to directions on the head. For the postcranial skeleton, we may also use cranial and caudal, which orient along the axis of the spine. For animals that don't carry their spine in an upright or vertical position, cranial and caudal will always denote the same directions.

    The vertebral axis is only one direction, and our bodies have two additional directions: front to back, and side to side. The terms for the front to back direction are:

    Anterior: Toward the front. The nose is on the anterior side of the head.

    Posterior: Toward the rear. The posterior side of the head is frequently covered in hair.

    Dorsal: In humans, toward the back of the torso. The shoulder blades are dorsal to the ribs.

    Ventral: In humans, toward the front of the torso. The navel is on the ventral aspect of the body.

    In humans, dorsal and ventral are mostly synonymous with posterior and anterior, and the latter terms are often used. In animals with habitual postures that are different than ours, dorsal and ventral retain an anatomical meaning that is unchanged and thus prevent confusion.

    Left and right are absolute terms instead of relative terms. These terms separate one half of the body from the other. The right arm will always be the right arm, and the right lung is right even though it is not as far right as the right arm.

    To refer to the position of a feature relative to another, the following terms are used:

    Medial: Closer to the midline, or dividing line between right and left halves, of the body. The neck is medial to the shoulder.

    Lateral: Farther from the midline. The eye is lateral to the nose.

    The limbs are special cases, because they can move a great deal relative to the spine. For the limbs, anterior, posterior, medial, and lateral are all relative terms used in reference to a particular limb position, called the anatomical position. For humans, the arms are in anatomical position when hanging at the sides of the body, palms forward, and the legs are in anatomical position in a normal standing posture, feet side by side. This means that the pinky side of the wrist is medial, and the thumb side is lateral. Superior and inferior are not used for the limbs at all. These terms are replaced by:

    Proximal: Closer to the point of attachment with the torso. In other words, closer to the shoulder or the hip. The elbow is proximal to the wrist.

    Distal: Farther from the point of attachment. The ankle is distal to the knee.

    Figure illustrating anatomical directions

    These terms can be somewhat confusing to learn, but they prevent a great deal of confusion in referring to bones and their features. The most common ones
    in this course will be anterior, posterior, superior, inferior, medial, lateral, proximal, and distal. Teeth and the hands and feet each have a few special directional terms, which will be introduced along with these anatomical areas.

  • Human and ape feet

    Mon, 2012-01-23 09:01 -- John Hawks
    Synopsis: 
    Laboratory exercise on the anatomical differences between human and ape toes

    At this station, you'll find some articulated human feet. "Articulated" means that the bones are assembled together at their joints -- two bones that articulate with each other are connected at a joint.

    You will also find some feet from two living species of great apes, gorillas and orangutans. These feet are obviously different from human feet in several respects. Most obviously, ape feet have an opposable first toe. The first toe in humans is often called the "big toe" or "great toe", but in these apes it is quite a bit shorter than the other toes. In anatomical terms, the first toe is called the hallux, and it is on the medial side of the foot, the one closest to the midline of the body.

    The other toes, which are lateral to the hallux, are substantially longer in apes than in humans. Each of these toes consists of three bones, which are called phalanges. The closest to the rest of the foot is the proximal phalanx, the furthest is the distal phalanx. The one in the middle of the toe is called the intermediate phalanx.

    Take a look at the intermediate phalanges in the human and ape feet. What do you notice about them? This is a substantial difference in the anatomy of human and ape feet, as human toes have greatly reduced all the phalanges but particularly the intermediate ones.

    Think of a hypothesis to account for the shorter toes in humans. Why would it make a difference how long the lateral toes are?

  • Canines

    Mon, 2011-10-17 23:27 -- John Hawks
    Synopsis: 
    Lab exercise introducing canines, including sizes of maxillary canines in hominoids.

    The canine teeth in humans range from pointy-shaped to incisor-like in shape. There is only one canine in each quadrant, and it is the third tooth just distal to the incisors.

    Upper canines are often denoted UC and lower canines are then LC (so that the left lower canine is LLC.

    In many other primates, the canine teeth project out far beyond the others. There is often a large space, or \term{diastema} between the upper canine and the lateral incisor. For many species, the canine teeth are the largest difference between male and female skulls.

    This station has the skulls of several kinds of primates. Measure the height of the right canine tooth in each maxillary dentition. This measurement is taken from the tip of the canine to the base of its enamel.

    Then measure the breadth of the first molar.

    Your assignment is to make a plot showing how canine height relates to molar breadth in this sample of primates. Are there any outliers in your plot?

    Are there any other features of the mandibles that seem to correlate with canine breadth?

  • Molars

    Tue, 2011-10-11 08:01 -- John Hawks
    Synopsis: 
    Laboratory exercise to introduce the terminology and anatomy of the molars.

    The most distal teeth are molars. Most humans have three molars, but many — especially in America — have their third molars (called wisdom teeth) extracted. Some people do not develop third molars at all, or they never erupt into occlusion. Molars have three or more cusps, and are used as grinding teeth.

    The upper molars are typically labeled with superscript numbers M1, M2, and M3, the lowers with subscript numbers M1, M2 and M3. Hence, the left lower first molar becomes LM1.

    Teeth have different directional terminology, referring specifically to the tooth row and the mouth. The direction toward the center front of the tooth row is mesial, and toward the rear of the tooth row is distal. For molars and premolars, the direction toward the cheek is buccal, and in toward the tongue is lingual.

    The two incisors, one canine, two premolars and three molars on both top and bottom are called the human dental formula. We write a dental formula as follows:

    2 1 2 3
    ___________
    2 1 2 3
  • Premolars

    Tue, 2011-10-11 07:37 -- John Hawks
    Synopsis: 
    Laboratory exercise to familiarize students with premolars in the dentition.

    The premolars are directly distal to (behind) the canines. Generally there are two premolars in each quadrant. Counting backward from the front of the jaw, there are normally two adult incisors, one canine, and then the two premolars, making them the fourth and fifth teeth in each row.

    In anthropology, we number these teeth differently from dentists, because the premolars in humans are homologous with the distal premolars in other mammals. So the human premolars are called the third and fourth premolars, even though we have only two of them! The lowers are numbered P3 and P4; the uppers P3 and P4.

    Premolars usually have two distinct points, or cusps, on their occlusal surface. In humans the lower third premolar is sometimes shaped very much like a canine tooth with only one cusp. In many primates, the P3 has only one large cusp that cuts against the distal edge of the upper canine, like a scissors.

    Examine the premolars at this station, both upper and lower. Learn to distinguish these from the other teeth.

    The two incisors, one canine, two premolars and three molars on both top and bottom are called the human dental formula. We write a dental formula as follows:

    2 1 2 3
    ___________
    2 1 2 3
  • Radius and ulna

    Mon, 2011-09-26 10:49 -- John Hawks
    Synopsis: 
    A lab exercise to learn the anatomy of the bones of the lower arm.

    The radius and ulna are the two bones of the lower arm. Rotation of the wrist is actually accomplished by a rotation of the radius around the ulna. The radius is on the lateral side of the arm, while the ulna is medial.

    The radius can turn at the elbow joint, and therefore the part of the radius that articulates with the humerus, called the radial head, has a rounded circumference that allows it to rotate in contact with the proximal ulna. The radius has a large tuberosity on the front, toward the medial side.

    The proximal ulna has a large notch bounded by two bony processes. The olecranon process fits into the olecranon fossa of the humerus, while the coronoid process fits against the anterior surface of the humerus when the elbow is flexed. Between these two processes, a half-moon shaped notch, called the semilunar or trochlear notch, fits strongly around the trochlea of the humerus, creating a stable and strong hinge joint to counter the weak but rotating joint of the radius. The proximal ulna has a notch for the radial head, called the radial notch, which is on the lateral side of the ulna.

    At the distal end, most of the proximal wrist joint is occupied by the distal radius, with the distal ulna free to rotate relative to the wrist. Both bones have pointed styloid processes, which extend on the medial and lateral sides of the wrist.

    What to do: At this station are many right and left radii and ulnae, including some fragmentary bones. Use your time to learn well how to tell right and left apart.

  • Humerus

    Mon, 2011-09-26 09:46 -- John Hawks
    Synopsis: 
    A lab introduction to the anatomy of the humerus.

    The bone of the upper arm is called the humerus. It articulates with the scapula at the shoulder joint, and the radius and ulna at the elbow.

    The proximal end of the humerus is dominated by a half-spherical articular surface, called the head, that forms the ball of the ball-and-socket joint of the shoulder. The head points medially into the shoulder joint. On the lateral side, a bump called the greater tubercle projects proximally.

    The distal end of the humerus has two articular surfaces. The first of these, called the trochlea, is a pulley-shaped surface that accommodates the ulna. The other, called the capitulum, is a small spherical structure lateral to the trochlea that articulates with the head of the radius. The capitulum is on the lateral side, the trochlea is medial.

    On the posterior surface, above the trochlea is a large dent, called the olecranon fossa. The proximal end of the ulna fits into this fossa when the elbow is extended.

    What to do: At this station are many right and left humeri, including some fragmentary bones. Work on telling right and left humeri from each other. You will find the distal end of the bone very helpful, with the trochlea medial and capitulum lateral, and the olecranon fossa on the posterior aspect.

  • Tibia and fibula

    Tue, 2011-09-06 01:49 -- John Hawks
    Synopsis: 
    A laboratory exercise to learn the basic anatomy of the lower leg and to side right and left tibiae.

    The tibia and fibula are the bones of the lower leg. The fibula forms the lateral part of the ankle joint, preventing dislocation in that direction.

    The tibia is much larger and thicker than the fibula. On its proximal end, the tibia has two condyles that provide a platform on which the distal condyles of the femur sit during weight support. The anterior surface of the tibia lies just under the skin, and is often called the shin. At the proximal end of the anterior surface is the large tibial tuberosity, which most people can feel just below their kneecap.

    The distal end of the tibia makes up much of the ankle joint, and the tibia has a distal projection on its medial side, called the medial malleolus, that can be felt on the inside surface of the ankle, and stabilizes the ankle in the medial direction.

    The tibial tuberosity is on the front, or anterior aspect, of the tibia, and the medial malleolus is medial. These two features are good guides to determining whether a tibia is from the right or left side. Remember, it is the skeleton's right or left, not yours as you look at the bone.

    Study questions: 
    1. At this station are many right and left tibiae and fibulae, including some fragmentary bones. See if you can determine which are right and which are left.
  • Femur

    Fri, 2011-09-02 01:24 -- John Hawks
    Synopsis: 
    A lab to introduce the anatomy of the femur.

    The femur is the bone of the upper leg. The proximal end of the femur connects to the hip joint. It is marked by a spherical ball, called the femoral head, that fits into the socket of the hip joint, the acetabulum. The head is connected to the shaft of the femur by an elongated segment of bone, called the femoral neck. Lateral to the neck, a large projection juts proximally off the top of the bone, called the greater trochanter. A smaller projection on the posterior surface of the femur, just below the neck, is called the lesser trochanter.

    The shaft of the femur is thick, and may be quite straight or slightly curved from front to back.

    The distal end of the femur connects to the knee joint. It is marked by two large articular processes, called condyles, which sit on top of the tibia.

    Telling a right femur from a left is easy if you know what to look for. The head of the femur connects to the hip joint, so it is toward the middle of the body. Using anatomical terminology, it's medial and proximal. The greater trochanter is on the lateral side of the bone, away from the middle of the body. The front side of the femur (called the anterior side) is fairly smooth. The back side (called the posterior side) has the lesser trochanter and the condyles both projecting back.

    If you align the femur so that the condyles and lesser trochanter are pointing backward, the head must point toward the hip. Also, if you place the condyles of a human femur flat on a table surface, the shaft of the bone will have an angle, called the valgus angle. It angles outward toward the hip from the center of mass of the body. So a right femur angles toward the right, a left femur angles toward the left.

    Remember, right and left refer to the skeleton's body, not the way you are looking at it!

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