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paleoanthropology, genetics and evolution

Homo erectus

  • Mandibles of early Homo and robust australopithecines

    Mon, 2012-11-12 22:36 -- John Hawks
    Synopsis: 
    A lab showing the variation of mandibles in early members of our genus.

    For anthropologists, Africa was a point of exceptional diversity between 2 million and 1.5 million years ago. In both East and South Africa, the fossil record presents evidence of several different hominin species. Some fossils belong to our own genus, Homo, and others belong to robust australopithecines.

    These two forms seem like they should be easy to tell apart. Robust australopithecines had extraordinarily large mandibles compared to living humans. Consider:

    • The main part of the mandible, which holds the teeth, is called the mandibular corpus. In robust australopithecines, this is often extremely thick and tall, with a large distance from the inferior border of the mandible to the teeth.
    • The portion of the mandible that extends upward to articulate with the temporal bone is called the mandibular ramus -- with one on both left and right sides. The mandibular ramus of many robust australopithecines is exceedingly tall, reflecting the very vertically tall faces of these hominins.
    • Robust australopithecines have hugely expanded premolars and molars, and greatly reduced incisors and canines. Early Homo has overall larger teeth than in living humans, but the proportions between the molars, premolars, incisors and canines is very much like people today.

    However, despite these obvious differences, the mandibles of early Homo and robust australopithecines are not always so easy to tell apart. This station has several mandibles from robust australopithecines, mainly from Australopithecus robustus from Swartkrans and Kromdraai, South Africa. There are also several mandibles of Homo erectus here, and a handful of mandibles that are likely early Homo but not definitely H. erectus.

    Can you tell them apart? Try seriating these from most humanlike to most robust australpithecine-like. Is there a clear dividing line between the two, or are there questionable specimens?

  • Meet Homo habilis

    Mon, 2011-11-07 23:44 -- John Hawks
    Synopsis: 
    A tour of four crania of Homo habilis

    This station has several of the key cranial specimens of Homo habilis, together with Sts 5, the representative of Australopithecus africanus. The H. habilis specimens include:

    • KNM-ER 1470, from Koobi Fora, Kenya, 1.9 million years old.
    • KNM-ER 1813, from Ileret, Kenya, 1.65 million years old.
    • OH 24, from Olduvai Gorge, Tanzania, 1.8 million years old.

    Another skull, KNM-ER 1805, is also included here. This skull may also represent H. habilis, or it may be something else.

    What to do: Examine the H. habilis crania compared to A. africanus and early Homo erectus. What makes these skulls more like Homo than Australopithecus?

    KNM-ER 1470 and KNM-ER 1813 are very different in size. Are they male and female of the same species, or do you think they are different species?

    What is KNM-ER 1805?

  • Statures of fossil Homo

    Tue, 2011-09-13 00:25 -- John Hawks
    Synopsis: 
    A laboratory exercise that applies regression equations to estimate the statures of some fossil hominin femora.

    Homo erectus and Neandertals were more or less human-sized. That may not be saying much, since we are so variable in stature ourselves.

    In this case, the fossils don't entirely speak for themselves. To estimate the sizes of ancient people, working with long bones, we must apply some kind of regression or other estimation method.

    1. KNM-ER 1481 is a complete femur from Koobi Fora, Kenya, approximately 1.9 million years old. Without any associated skull or teeth, we can't be sure what species it represents. Many scientists attribute it to early Homo because of its differences from known australopithecine femora.
    2. The Trinil femur was found by Eugene Dubois in 1892 as he excavated fossil beds at Trinil, Java. He had found a human skullcap the year before, and after finding the femur's humanlike anatomy, Dubois named a new species, Pithecanthropus erectus. This is the original Homo erectus femur. Today, we are less certain about its age and association with the partial skull. It may be a million years old, but it may be substantially younger.
    3. The femur from Spy, Belgium, represents a Neandertal who lived around 45,000 years ago. This femur is part of a more complete skeleton, and exhibits many of the characteristic features of Neandertal long bones, including the great thickness and curvature of the shaft and very large joint surfaces.
    4. What to do: Here you will examine the fossil cast femora, using regression equations to predict stature of the individual.

      1. Determine the sex of the individual. The femur head diameter is a relatively good indicator of sex. If it is less than 44 mm, the individual is likely to be a female. More than 46 mm, and the individual is likely to be a male. In between these values, you may need more information — either from the rest of the skeleton or from the size and robusticity of the femur itself.
      2. Measure the maximum length of the femur. This measurement is taken using the osteometric board, and represents the maximum distance from any points on the proximal and distal ends of the bone. Take your measurement in centimeters.
      3. Apply the correct regression equation. These are specific to sex and race. The femora at this station come from donated anatomy collections from the early 20th century, and represent people of European ancestry. The male and female regression equations for this population are listed at right.
    Study questions: 
    1. What are some weaknesses of estimating body size for fossil humans by applying a regression drawn from a contemporary human population?
Subscribe to Homo erectus

Neandertals

For years, I've worked on their bones. Now I'm working on their genes. Read more about the science studying these ancient people.

Denisova

From a finger bone of an ancient human came the record of a completely unexpected population. My lab is working on the science of the Denisova genome.

Acceleration

The advent of agriculture caused natural selection to speed up greatly in humans. We're uncovering some of the ways that populations have rapidly changed during the last 10,000 years.

Malapa

Just outside Johannesburg, the Malapa site is producing some of the most exciting finds in human evolution. This site is the headquarters of the Malapa Soft Tissue Project.